Showing posts with label TFR. Show all posts
Showing posts with label TFR. Show all posts

Sunday, January 04, 2026

Pronatalism via Population Genetics

---

Japan’s Population in the Next Thousand Years: Fertility, Assortative Mating, and the Shape of the Future

Two years ago, I wrote about the likely dynamics of population size in advanced capitalist countries, using Japan as my example. People note that in advanced countries the TFR is well below replacement, and then conclude that the populations in these countries are on the road to extinction. 'More migration,' they cry, showing that these discussions are essentially ideological rather than scientific.

The misfortune of the human race, if we can call it that, is that we evolved - mostly - to like having sex rather than babies. For most of our evolutionary history one thing led to another, however, so this didn't matter in Darwinian terms. However, modern capitalism has provided both effective contraception and many economic and personal incentives to do more interesting things than change nappies and deal with troublesome toddlers. This disconnection between sex and reproduction has led to the collapse in TFR.

However, a minority of men and women actually do want to have families - their emotional drives prompt them to have children despite the many distractions. Evolutionary theory tells us, of course, that future populations are constructed from their ancestors who reproduced, indicating that the prevalence of alleles which promote child-bearing directly will tend to increase in the population, generation by generation. 

This is selective advantage in operation: over time the population will be replaced by individuals who actually want to raise families. Let's see how this would work out in Japan.

Japan currently stands near 123 million people, yet its total fertility rate (TFR) is ~1.3, far below the ~2.1 births per woman needed for replacement. What follows sketches thousand-year outcomes using simple demographic arithmetic and population-genetic reasoning.

1. Straight-line decline

With TFR ≈ 1.3 and no offsetting forces, each generation replaces ~60% of itself. Headcount falls steeply: a remnant population of only tens of millions by 2200 and potentially a group in the hundreds of thousands by 3000. Nevertheless, this is a cultural and institutional contraction rather than a biological extinction risk although it's hard to see an advanced civilisation surviving in anything like its current form.

2. Genetics and slow selection

Completed fertility has modest heritability (≈0.2). Directional selection for a stronger “baby-drive” nudges fertility up by roughly +0.01 to +0.03 births per woman per generation. Timescale to reach replacement by genetics under the assumption of random partner selection alone is centuries; the population would likely bottom at a few percent of today’s size before any rebound.

3. Immigration as a bridge

Immigration offsets natural decrease for 1–2 generations because empirically, migrants arrive young and initially have slightly higher fertility. Likely convergence to host norms and global low fertility limit this as a century-scale fix. It buys time; it does not determine the endpoint.

4. Assortative mating as the hidden accelerator

Key dynamic: if a minority of strongly pronatal men and women mostly partner with each other, their subpopulation grows multiplicatively while the mainstream population shrinks towards zero. 

Japan’s future is not locked to straight-line decline. Assortative mating among pronatal families can bend the curve upward in roughly a century, especially with supportive culture and policy. 

Immigration buys time; automation and biotech buffer the costs of small populations. The decisive variables are partner choice, norm transmission, and targeted pronatal policy. These matter more than slow genetic drift alone.

In fact with the pronatal population exhibiting a TFR of 3.0 (three child families on average) it would take less than 250 years for Japan's population to bounce back to its current 123 million people.

Thursday, January 11, 2024

Japan’s population will bounce back - sort of


From Wikipedia

Japan's TFR (Total Fertility Rate) is around 1.34. This is the average number of children a Japanese family will have in contemporary Japan. The TFR which maintains a steady population level is 2.1 (some fraction of children do not reproduce through death, ill-luck or disinclination).

People conclude that the Japanese as a country-community are doomed. Since European TFRs are similar, averaging around 1.6, this repugnant conclusion seems quite generalisable.

But people who write sensational stories in the media about this pay only lip service to evolutionary theory, they don't operationalise it. This bland, average TFR hides subpopulations, some of which don't reproduce at all, but others of which are quite prolific in family size.

Since there is parental genetic influence on the size of family [1] we can treat this fecundity-variance as setting up a process of genetic truncation selection. The structure and incentives of Japanese society (a proxy for almost all advanced countries) poses enormous Darwinian selection pressure against those individuals with genotypes which encourage them to fail to reproduce adequately.

---

A simple Fermi estimate (a back of the envelope calculation) allows us to predict the result of this massive selection pressure on the Japanese population, in generation time steps.

Simplifying the numbers in [2] below, divide Japanese families into four categories with the following endogenous TFRs:

 - category zero is 25% of the population with TFR zero

 - category one is 33% of the population with TFR one

 - category two is 33% of the population with TFR two

 - category three is 9% of the population with TFR three.

So this is the population structure at generation 0. 

Assume for ease of calculation that the population of Japan is 123 million. We divide this into three generational age cohorts: 0-25, 25-50,50-75 and assume each cohort is equally numerous at 41 million people.

Over one generation the upper cohort dies (losing 41 million people) and the middle cohort reproduces to replace them (or not).

To make things more intuitive we will replace the percentages with 100 notional families representing those individuals who reproduce, namely the 41 million strong middle cohort. Call them 40 million to make the sums easier. So each 'family' represents 400,000 people. 

What happens to the population as the generations roll over?

Generation 1: 2025 => 2050

We started with 123 million. The elderly cohort dies leaving 82 million. And now the logic of extreme selection kicks in: 

 - category 0, 25 families, are culled from the gene pool and vanish 

 - the 33 families of category 1 are replaced by 16

 - the category two families simply replace themselves, remaining 33

 - the category three families increase their number by 50% to fourteen.

Total number of families is 16 + 33 + 14 = 63. The newborn population is 63 * 400,000 = 25 million. The new population is 82 million + 25 million =  107 million. This is consistent as it happens with the chart at the head of this post.

(Note: the resulting TFR here is 25/20.5 = 1.2 so our back of the envelope calculation is a slight under-estimate).

Quite a drop from 125 million.

What happens next? Those alleles which support a propensity to have children have increased in frequency, but we know the heritability of this trait is not high. So as generation 1 transitions to generation 2 in 2075, we will still see some families having zero offspring (or never forming a family unit at all) while the percentage of those families favouring more offspring will increase.

The modelling would be complex and noisy, but the direction of travel is clear. Darwinian selection is doing its work and a new population being selected for, composed of those families interested in having children despite all the many modern distractions. 

Inexorably, the population of individuals who genetically underpin category three will rise exponentially and will eventually dominate in the population, which will itself substantially rise in numbers if it can manage to sustain itself.

There is an interesting implication, which escapes no-one who thinks about this in an informed way, that any other genetically-informed traits which co-vary with fertility-promoting alleles will also be selected for. I admire many aspects of Japanese culture but my ignorance of the details of Japanese society is pretty much total. Still, someone much more informed than me should take a look.

---

[1] Heritability of family size (from Bard)

Here are some examples of heritability estimates for family size in different countries:

Denmark: 15-20%

United States: 20-30%

Australia: 25-30%

Netherlands: 30-40%

Japan: 20%

-

[2] From Bard - family sizes in Japan

Here are some specific percentages you might find helpful, based on data from 2020:

Percentage of women aged 45-49 with no children: 22.7%

Percentage of women aged 45-49 with one child: 32.5%

Percentage of women aged 45-49 with two children: 34.2%

Percentage of women aged 45-49 with three or more children: 10.6%